Vitamin K promotes VKORC1L1-dependent endothelial cell regeneration by limiting ferroptosis and TNF-mediated inflammation

https://doi.org/10.1007/s00392-025-02625-4

Elena Repges (Bonn)1, A. Aksoy (Bonn)1, K. Czogalla-Nitsche (Bonn)2, J. Oldenburg (Bonn)2, C. F. H. Müller (Bonn)1, V. Tiyerili (Waldbröl)3, A. Zietzer (Bonn)1, F. Jansen (Köln)4, G. Nickenig (Bonn)1, S. Zimmer (Bonn)1, M. Al Zaidi (Bonn)1

1Universitätsklinikum Bonn Medizinische Klinik und Poliklinik II Bonn, Deutschland; 2Uniklinik Bonn Experimentelle Hämatologie und Transfusionsmedizin Bonn, Deutschland; 3Kreiskrankenhaus Waldbröl GmbH Klinik für Innere Medizin, Kardiologie Waldbröl, Deutschland; 4Gemeinschaftspraxis Kardiologie Köln am Neumarkt Köln, Deutschland

 

Background:

Vitamin K exerts anti-oxidative properties which were proposed to be mediated by the vitamin K cycle enzyme VKORC1L1 (vitamin-K-2,3-epoxide-reductase-complex-subunit-1-like-1). Anti-oxidative mechanisms are critical for proper endothelial function and to ensure regeneration of the endothelium upon injury. Here, we investigated the role of vitamin K on endothelial cell regeneration and anti-oxidation. 

 

Methods:

Human coronary artery endothelial cells (HCAEC) were treated with the vitamin K2 vitamer MK-7 (Menaquinone-7) to evaluate its effects on endothelial regeneration and anti-oxidative capacity. Cell viability was measured using the AlamarBlue assay, proliferation was assessed with EdU incorporation, apoptosis was evaluated via caspase 3/7 activity, and ferroptosis was determined by assessing viability following RSL3 stimulation. Lipid peroxidation was analyzed using Bodipy 581/591 staining, with detection by flow cytometry and immunofluorescence. Endothelial-to-mesenchymal transition and NF-κB/ICAM-1 signaling were assessed using qPCR and immunocytochemistry. RNA interference was applied to silence VKORC1L1 and VKORC1.

 

Results:

MK-7 treatment promoted a regenerative endothelial phenotype, marked by increased cell viability and proliferation and decreased apoptosis and ferroptosis. MK-7 also enhanced anti-oxidative capacity by reducing lipid peroxidation, ROS formation, endothelial-to-mesenchymal transition, and pro-inflammatory NF-κB/ICAM-1 signaling. Knockdown of VKORC1L1, but not of the canonical VKORC1, led to increased oxidative stress and inflammation. Importantly, MK-7 required VKORC1L1, but not VKORC1, to inhibit lipid peroxidation, ROS formation, and NF-κB activation. RNA sequencing revealed that VKORC1L1 silencing upregulated TNF expression. Accordingly, VKORC1L1-deficient endothelial cells showed elevated TNF secretion under pro-inflammatory conditions when compared to VKORC1-deficient or control cells. TNF secretion in VKORC1L1-silenced cells was driven by NF-κB signaling and was dependent on lipid peroxidation, which was effectively inhibited by Ferrostatin-1. Finally, blocking TNF shedding with TAPI-1 reduced ROS formation and inflammation in VKORC1L1-silenced HCAEC.

 

Conclusion: 

Vitamin K promotes endothelial cell regeneration by mitigating lipid peroxidation, ferroptotic cell death, and TNF-mediated inflammation. These protective effects are mediated by VKORC1L1 but not the canonical vitamin K enzyme VKORC1.



 

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